基金Acknowledgements We are grateful to S.A. Shilova who encouraged and supported the study. We thank L.E. Savinetskaya, V.S. Popov, N.S. Vasiliev, I.A. Volodin, E.V. Volodina, V.A. Matrosova, A.A. Kochetkova and other members of our field team, and thank Dr. Michael Birman for polishing the revised paper. Comments and suggestions of two anonymous reviewers to a previous version of this manuscript were very helpful. The study was financially supported by the Russian Foundation for Basic Research （grants 10-04-01304a and 12-04-31279）.